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Santalum paniculatum

Hook. & Arn. (1832)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Santalopsida Order: Santalales Family: Santalaceae Genus: Santalum

‘iliahi [iliahi], sandalwood

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Description

Key Characters:

Growth Form: Shrubs or trees 3–10(–20) m tall. Apparently hemiparasitic on roots of other plants.

Stems:

Roots:

Leaves: Leaves simple. Opposite or rarely alternate. Blades ovate, elliptic, or obovate, (2.5–)5.5–6(–8) cm long, (2–)2.54.5 cm wide. Upper surfaces glossy; lower surfaces dull and ± pale, often glaucous, brittle; evergreen, simple, ± conspicuously discolorous, sometimes yellowish orange to bluish or olive green. Margins entire. Petioles 2–15 mm long. Stipules absent.

Flowers: Flowers in terminal and ± axillary, compound cymes. Flowers bisexual (perfect), with a sweet fragrance, greenish in bud, about as long as wide. Calyx (sepals) absent. Corolla greenish, tinged with brown, orange, or salmon after opening, 4– 8 mm long. Stamens 4(5). Ovary inferior; stigma 2–4-lobed.

Fruit: Drupes purple to black at maturity; 10–12 mm long; with an apical receptacular ring. Seeds muricate; with pointed beak on one end.

Ploidy: 2n = 40*

Habitat: Scattered in dry woodland on lava or on cinder cones; to higher elevation wet forest or secondary <i>Metrosideros</i> forest.

Elevation Range: 450–2,000 (–2,550) m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

O'ahu Only found in cultivation
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Bot. Beechey Voy.: 94 (1832)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Santalum paniculatum Kau-Kona near boundary Kahuku Ranch, Kipuka between two 1907 flows from Mauna Loa Preserved_Specimen Glover, J.W. Hawaii BISH 1947-06-23
2 Santalum paniculatum Mauna Kea; east of Hale Pohaku Camp Preserved_Specimen Adee, K.T. Hawaii BISH
3 Santalum paniculatum forest of Ainahau near Kilauea Preserved_Specimen Rock, J.F.C. Hawaii BISH 1958-10-06
4 Santalum paniculatum Kona, uplands of Puu Lehua, Greenwell Ranch Preserved_Specimen Rock, J.F.C. Hawaii BISH 1958-04-24
5 Santalum paniculatum S. slope of Hualalai Preserved_Specimen Degener, O. Hawaii BISH 1967-07-27
6 Santalum paniculatum Kilauea crater road Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
7 Santalum paniculatum Mauna Kea Preserved_Specimen Bryan, L.W. Hawaii BISH 1951-01-19
8 Santalum paniculatum Kilauea Preserved_Specimen Hillebrand, W.B. Hawaii BISH
9 Santalum paniculatum Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
10 Santalum paniculatum Kau Desert, Haw. Natl. Park Preserved_Specimen Russ, G.W. Hawaii BISH 1927-04-01
11 Santalum paniculatum Hawaii Preserved_Specimen Neal, M.C. Hawaii BISH 1958-01-01
12 Santalum paniculatum unknown Preserved_Specimen Hawaii BISH
13 Santalum paniculatum N.Kona, Kaupulehu, dry weedy lava slopes 2 mi east of Huehue Preserved_Specimen Webster, G.L. 1822 Hawaii BISH 1948-06-23
14 Santalum paniculatum Kona Preserved_Specimen Hillebrand, W.B. Hawaii BISH
15 Santalum paniculatum Flats Southeast of Waha Pele. Keahou no. 2 Hualalai, No. Kona Preserved_Specimen Fosberg, F.R. 41644 Hawaii BISH 1961-03-22
16 Santalum paniculatum crater rim trail betw. Kialuea Military Camp and Volcano Observatory, Hawaii National Park Preserved_Specimen Fosberg, F.R. 41780 Hawaii BISH 1961-03-27
17 Santalum paniculatum Between Waha Pele and Puu Ikaaka, Hualalai, N. Kona Preserved_Specimen Fosberg, F.R. 41645 Hawaii BISH 1961-03-22
18 Santalum paniculatum Hanehane, Kona Preserved_Specimen Forbes, C.N. 175 Hawaii BISH 1911-06-16
19 Santalum paniculatum Puuoikaaka Crater. slops of Mt. Hualalai Preserved_Specimen Rock, J.F.C. 10048 Hawaii BISH 1912-02-13
20 Santalum paniculatum unknown Preserved_Specimen BISH
21 Santalum paniculatum E of Hale Pohaku, Mauna Kea arid lava ledges Preserved_Specimen Degener, O. 19970 Hawaii BISH 1949-09-04
22 Santalum paniculatum Puu Huluhulu cinder cone Preserved_Specimen Degener, O. 19984 Hawaii BISH 1949-08-20
23 Santalum paniculatum N. Kona, SW slopes of Puu Hualalei, NW of Waha Pele Volcanic Cone `a`a lava, scattered trees and shrubs Preserved_Specimen Stauffer, H.U. 5898 Hawaii BISH 1964-04-19
24 Santalum paniculatum Hilo, Olaa-Hiulani old lava flow Preserved_Specimen Skottsberg, C.J.F. 429 Hawaii BISH 1922-09-08
25 Santalum paniculatum Between Huehue and Puuwaawaa dry lava fields Preserved_Specimen Rock, J.F.C. 3728 Hawaii BISH 1909-06-01
26 Santalum paniculatum Northwest of Mauna Loa Boys School, Kulani Project, South Hilo District A small stand of 3 trees and 1 sapling. Growing in an open Metrosideros forest with a Styphelia scrub with a Deschampia grassland. Pahoehoe substrate. Preserved_Specimen Clarke, G. 565 Hawaii BISH 1981-02-12
27 Santalum paniculatum Stainback Hwy wind, sun, cold, lava Preserved_Specimen Degener, O. 21850 Hawaii BISH 1952-02-15
28 Santalum paniculatum Kona, Kikiaeae with koa, in open stands Preserved_Specimen Christ, J.H. Hawaii BISH 1958-05-16
29 Santalum paniculatum N. Kona, SW slopes of Puu Hualalei, NW of Waha Pele Volcanic Cone volcanic ash with open low shrubby vegetation, with Dodonaea, Metrosideros, Wikstroemia, Exocarpos Preserved_Specimen Stauffer, H.U. 5900 Hawaii BISH 1964-04-19
30 Santalum paniculatum Bernice Pauahi Bishop Museum Native Garden. Cultivated in BPBM Native Hawaiian Garden. With Ipomoea batatas, Plumbago zeylanica, Pipturus albidus, and Acacia koaia. Preserved_Specimen Walvoord, N. 2023010 Oahu BISH 2023-02-08
31 Santalum paniculatum 19 mi from Waimea toward Kona aa desert Preserved_Specimen Degener, O. 5295 Hawaii BISH 1926-08-18
32 Santalum paniculatum Between Puuwaawaa and Waimea aa desert Preserved_Specimen Degener, O. 5291 Hawaii BISH 1926-08-18
33 Santalum paniculatum Royal Gardens, Kalapana, ca. 100 ft. off Prince St. towards Hawaii Volcanoes National Park Closely associated with Canthium, Diospyros, Metrosideros, Dodonaea, and Christmas berry. Very low elevation. Substrate `a`a. Preserved_Specimen Davis, J. 1004 Hawaii BISH 1985-05-24
34 Santalum paniculatum Kawaihae, DHHL Survey, Supp. Sta. P Pasture above kiawe forest with Prosopis, Leucaena leucocephala, Opuntia ficus-indica, Solanum sodomeum. Preserved_Specimen Perlman, S. 10289 Hawaii BISH 1989-08-04
35 Santalum paniculatum N. Kona, SW slopes of Puu Halalei, Keahou `a`a lava, pasture Preserved_Specimen Stauffer, H.U. 5895 Hawaii BISH 1964-04-19
36 Santalum paniculatum Pohakuloa Training Area near Puu Koko, mauka of pole 116 alluvum of small gulch Preserved_Specimen Stemmermann, R.L. 675 Hawaii BISH 1975-01-08
37 Santalum paniculatum Between Huehue and Puuwaawaa dry lava fields Preserved_Specimen Rock, J.F.C. 3607 Hawaii BISH 1909-06-01
38 Santalum paniculatum Puu Laau exclosure, Mauna Kea Sophora forest Preserved_Specimen Stemmermann, R.L. 680 Hawaii BISH 1975-01-08
39 Santalum paniculatum Haloa Aina reforestation Project site. Hawaii State TMK 379001014. Planted within 5m of the Haloa Aina Greenhouse Manicured lawn setting. The area is regularly mowed and temperature ranges from 26deg C to 1.6deg C. Other plants: Acacia koa, Sophora chrysophylla, Pitosporum spp, Pipturus albidus, and Myoporum sandwicense Preserved_Specimen Speetjens, T.M. Hawaii BISH 2023-01-23
40 Santalum paniculatum NW of Kaalualu dry aa forest Preserved_Specimen Degener, O. 5318 Hawaii BISH 1929-09-13
41 Santalum paniculatum Kapapala Forest Reserve, Kau District, growing beside jeep road Growing in open koa-'ohi'a parkland with native shrubs and grasses. Preserved_Specimen Cuddihy, L.W. 812 Hawaii BISH 1981-07-02
42 Santalum paniculatum N. Kona, Puuanahulu dry forest on aa; with Myoporum sandwicense st.johnii Preserved_Specimen Webster, G.L. 14006 Hawaii BISH 1968-06-30
43 Santalum paniculatum N. Kona, Puuwaawaa lava flows Preserved_Specimen Rock, J.F.C. 25717 Hawaii BISH 1957-03-14
44 Santalum paniculatum N.Kona, Puuwaawaa Ranch in Acacia koa, pasture Preserved_Specimen Herbst, D.R. 5481 Hawaii BISH 1975-09-24
45 Santalum paniculatum SW flank of Mauna Loa, Manuka open Metrosideros forest on unweathered aa with scrub; with Polypodium, Coprosma, Dodonaea, Vaccinium Preserved_Specimen Higashino, P.K. 6247 Hawaii BISH 1977-07-22
46 Santalum paniculatum Kapua, makai of Hwy Diospyros forest Preserved_Specimen Stemmermann, R.L. 694 Hawaii BISH 1975-01-10
47 Santalum paniculatum NE of Kaalualu Dry aa forest. Preserved_Specimen Degener, O. 5319 Hawaii BISH 1929-09-13
48 Santalum paniculatum Kau desert Preserved_Specimen Russ, G.W. Hawaii BISH 1929-04-01
49 Santalum paniculatum Haloa Aina reforestation Project site. Hawaii State TMK 379001014. Planted within 5m of the Haloa Aina Greenhouse Manicured lawn setting. The area is regularly mowed and temperature ranges from 26deg C to 1.6deg C. Other plants: Acacia koa, Sophora chrysophylla, Pitosporum spp, Pipturus albidus, and Myoporum sandwicense Preserved_Specimen Speetjens, T.M. Hawaii BISH 2023-01-23
50 Santalum paniculatum near Kilauea MC, National Park Metrosideros forest on pahoehoe lava; in small depression Preserved_Specimen Takeuchi, W.N. 11 Hawaii BISH 1984-05-12
51 Santalum paniculatum 2.5 mi SSE of Puu Kapele Naio, Aalii Preserved_Specimen Fagerlund, G.O. 529 Hawaii BISH 1943-04-16
52 Santalum paniculatum Mauna Kea, SW slope above military camp, Puu Laau volcanic sand, thicket of sandalwood with grasses Preserved_Specimen Stauffer, H.U. 5906 Hawaii BISH 1964-04-21
53 Santalum paniculatum saddle between Mauna Loa and Mauna Kea volcanic hills; sand and rubble; open shrubby vegetation with Myoporum, Dodonaea and grasses Preserved_Specimen Stauffer, H.U. 5905 Hawaii BISH 1964-04-21
54 Santalum paniculatum Between Waha Pele and Puu Ikaaka, Hualalai, N. Kona tall shrub Preserved_Specimen Fosberg, F.R. 41645 Hawaii BISH 1961-03-22
55 Santalum paniculatum S slope of Hualalai decadent Sophora forest, pasture and grass Preserved_Specimen Degener, O. 31344 Hawaii BISH 1967-07-27
56 Santalum paniculatum Keauhou 2 Ahupuaa, W slope of Mauna Loa, heading WNW from ca. 1.3 km N of Kohalapilau toward Highway 11. Transect 61, Station 45. Dry koa-mamane community with an exotic grass and scattered native shrub understory. Preserved_Specimen Jacobi, J.D. 1042 Hawaii BISH 1978-07-01
57 Santalum paniculatum 2 mi NE of Huehue Ranch House, Kaupulehu, N. Kona in scrub on aa lava Preserved_Specimen St.John, H. 22451 Hawaii BISH 1946-12-28
58 Santalum paniculatum Saddle Rd., small side road leading mauka from pole 116 alluvial fan from Mauna Kea with Chenopodium, Sophora, Euphorbia, Myoporum, Santalum, Dodonaea Preserved_Specimen Stemmermann, R.L. 678 Hawaii BISH 1975-01-08
59 Santalum paniculatum Keauhou Ranch On top of aa flow ridge. Native scrub and open Metrosideros forest. Preserved_Specimen Davis, J. 216 Hawaii BISH 1980-02-05
60 Santalum paniculatum National Park, Kipuka Puaulu between fenced area and picnic area closed canopy ohia-koa forest and exotic grass understory Preserved_Specimen Stemmermann, R.L. 667 Hawaii BISH 1974-11-01
61 Santalum paniculatum Kaupulehu Forest S of Puuwaawaa, N. Kona rough aa lava, dry forest; with Diospyros ferrea sandwicensis, Colubrina, Nototrichium, Kokia, Canthium, Alphitonia Preserved_Specimen Carlquist, S. 2102 Hawaii BISH 1966-07-31
62 Santalum paniculatum Kilauea, between Volcano House and military camp volcanic ash and pahoehoe lave, shrubby forest Preserved_Specimen Stauffer, H.U. 5903 Hawaii BISH 1964-04-20
63 Santalum paniculatum upslope of Kalapana Reynoldsia, Canthium, Schinus, Wikstroemia, Bobea Preserved_Specimen Stemmermann, R.L. 662 Hawaii BISH 1974-11-01
64 Santalum paniculatum mauka of pole 116, near Puu Koko, Pahakuloa Training area, W of headquarters alluvum of small gulch Preserved_Specimen Stemmermann, R.L. 673 Hawaii BISH 1975-01-08
65 Santalum paniculatum Puu Huluhulu, Humuula, N. Hilo hill kipuka, open woods Preserved_Specimen St.John, H. 23932 Hawaii BISH 1949-12-24
66 Santalum paniculatum 2 mi NE of Huehue Ranch House, Kaupulehu, N. Kona in scrub on aa lava Preserved_Specimen St.John, H. 22452 Hawaii BISH 1946-12-28
67 Santalum paniculatum makai of Puu Ohohia, Mauna Loa kipuka Preserved_Specimen Degener, O. 31382 Hawaii BISH 1966-11-29
68 Santalum paniculatum 3 mi W of Volcano House, National Park with Coprosma ernodeoides, Dodonaea, Vaccinium, Metrosideros; greyish volcanic soil Preserved_Specimen Henrickson, J.S. 4069 Hawaii BISH 1969-08-06
69 Santalum paniculatum 4 km S of benchmark 2487, Volcano Rd; 4-6 km S of Puu Kamoamoa wet ohia forest, native trees and shrub, treefern; semi shaded Preserved_Specimen Jacobi, J.D. 1341 Hawaii BISH 1979-06-07
70 Santalum paniculatum Southwest flank of Mauna Loa, in the ahupuaa of Puu Ohohia semi closed Metrosideros and Acacia forest, understory Dodonaea, Styphelia, Vaccinium on cinder. Preserved_Specimen Higashino, P.K. 6315 Hawaii BISH 1977-08-05
71 Santalum paniculatum Waimea Kona Rd., opposite Puuwaawaa Dry, open forest on`a`a lava Preserved_Specimen Fosberg, F.R. 41692 Hawaii BISH 1961-03-23
72 Santalum paniculatum N. Kona, Kaupulehu grassy lava flow with scattered trees Preserved_Specimen Stauffer, H.U. 5907 Hawaii BISH 1964-04-21
73 Santalum paniculatum Kau near 1920 flow Preserved_Specimen Skottsberg, C.J.F. 1880 Hawaii BISH 1926-09-20
74 Santalum paniculatum Kilauea crater below Volcano House ledges along trail Preserved_Specimen Skottsberg, C.J.F. 3381 Hawaii BISH 1938-09-18
75 Santalum paniculatum National Park, Kipuka Puaulu between fenced area and picnic area closed canopy ohia-koa forest and exotic grass understory Preserved_Specimen Stemmermann, R.L. 669 Hawaii BISH 1974-11-01
76 Santalum paniculatum South Kona District: SW slopes of Mauna Loa, Kapua `a`a lava with scattered trees, shrubs, ferns Preserved_Specimen Stauffer, H.U. 5902 Hawaii BISH 1964-04-20
77 Santalum paniculatum Along a road leading to Kiholo, Kona Coast growing on a'a lava with Erythrina, Metrosideros, Xylosma, Lantana. Preserved_Specimen Herbst, D.R. 1588 Hawaii BISH 1970-02-22
78 Santalum paniculatum N. Kona, SW slopes of Puu Hualalei, NW of Waha Pele Volcanic Cone Steep inside slope of cone, Aa-lava, with scattered Metrosideros and Dodonaea Preserved_Specimen Stauffer, H.U. 5896 Hawaii BISH 1964-04-19
79 Santalum paniculatum Kaohe Game Management area just .5 mile north of Ahumoa on makai side of a jeep road Growing in an open Sophora-Myoporum scrub forest. Preserved_Specimen Clarke, G. 271 Hawaii BISH 1979-10-09
80 Santalum paniculatum 20 mi from Waimea toward Kona aa desert Preserved_Specimen Degener, O. 5292 Hawaii BISH 1926-08-20
81 Santalum paniculatum Northwest of Mauna Loa Boys School, Kulani Project, South Hilo District A small stand of 3 trees and 1 sapling. Growing in an open Metrosideros forest with a Styphelia scrub on a pahoehoe substrate. Preserved_Specimen Clarke, G. 566 Hawaii BISH 1981-02-12
82 Santalum paniculatum Puuwaawaa, rough lava Preserved_Specimen MacDaniels, L.H. 303 Hawaii BISH 1926-12-01
83 Santalum paniculatum SE of Kauunuoku, Hualalai, N. Kona Occasional in scrub on lapilli ash. Preserved_Specimen Fosberg, F.R. 41651 Hawaii BISH 1961-03-22
84 Santalum paniculatum Saddle Rd., Humuula, S slope of cinder cone, 1 mi S of Oonaokoili, N. Hilo sparse upper forest Preserved_Specimen St.John, H. 22378 Hawaii BISH 1946-12-26
85 Santalum paniculatum Norman Greenwell property. Near Puu Lehua Hualalai Present dominant in plain. Kikuyu grass, Myoporum, Sophora, some koa. Preserved_Specimen Stemmermann, R.L. 6915 Hawaii BISH 1984-09-23
86 Santalum paniculatum Keauhou Ranch, Kau District Growing in disturbed open Metrosideros, with few Acacia koa; shrubs Dodonaea, Styphelia and pasture of Eragrostis brownei, Setaria geniculata and Sacciolepis indica Preserved_Specimen Davis, J. 402 Hawaii BISH 1980-10-16
87 Santalum paniculatum Manuka Mauka In wooded Ripuka Preserved_Specimen St.John, H. 11342 Hawaii BISH 1931-12-26
88 Santalum paniculatum S. Kona among lava flows Preserved_Specimen Rock, J.F.C. Hawaii BISH 1958-06-01
89 Santalum paniculatum East of Hale Pohaku, Mauna Kea In arid Edwardsia forest. Preserved_Specimen Degener, O. 19971 Hawaii BISH 1949-09-04
90 Santalum paniculatum N. Kona, SW slopes of Puu Hualalei, NW of Waha Pele Volcanic Cone `a`a lava, scattered trees and shrubs Preserved_Specimen Stauffer, H.U. 5897 Hawaii BISH 1964-04-19
91 Santalum paniculatum SE of Kauunuoku, Hualalai, N. Kona low shrub Preserved_Specimen Fosberg, F.R. 41655 Hawaii BISH 1961-03-22
92 Santalum paniculatum Kilauea, Chain of Craters Rd., 0.8 km SE of Makaopuhi Crater dense forest on `a`a lava, with Metrosideros, Ilex, many ferns Preserved_Specimen Stauffer, H.U. 5904 Hawaii BISH 1964-04-20
93 Santalum paniculatum Kaupulehu FR W of lava flow, fenced Preserved_Specimen Wagner, W.L. 4902 Hawaii BISH 1983-07-28
94 Santalum paniculatum Saddle Road, 23.5 mi from Hilo kipuka Preserved_Specimen Degener, O. 20205 Hawaii BISH 1949-08-26
95 Santalum paniculatum makai of Ocean View Estates, Kau mixed ohia forest on aa Preserved_Specimen Stemmermann, R.L. 6664 Hawaii BISH 1980-06-06
96 Santalum paniculatum S. Kohala District, near Puu Heewai, ca. 2 miles SE of Keamuku Growing on edge of small gully. Preserved_Specimen Davis, J. 655 Hawaii BISH 1981-12-03
97 Santalum paniculatum Kau district, Keauhou Ranch Growing under Metrosideros tree in open Metrosideros scrub on pahoehoe substrate. Preserved_Specimen Cuddihy, L.W. 172 Hawaii BISH 1980-01-31
98 Santalum paniculatum Kahulea, 6.1 mi from Kalae turnoff kipuka in 1907 flow Preserved_Specimen Ozaki, E.T. 1302 Hawaii BISH 1954-12-23
99 Santalum paniculatum Kaupulehu Forest Reserve. Adjacent to the Kaupuleho Lava Flow. Dry forest, part of akipuka, growing adjacent to the Kaupuleho Lava Flow; aa lava. Preserved_Specimen Stern, W.L. 3121 Hawaii BISH 1974-03-28
100 Santalum paniculatum Haloa Aina reforestation Project site. Hawaii State TMK 379001014. Planted within 5m of the Haloa Aina Greenhouse Manicured lawn setting. The area is regularly mowed and temperature ranges from 26deg C to 1.6deg C. Other plants: Acacia koa, Sophora chrysophylla, Pitosporum spp, Pipturus albidus, and Myoporum sandwicense Preserved_Specimen Speetjens, T.M. Hawaii BISH 2023-01-23
101 Santalum paniculatum Puu Huluhulu, saddle road cinder cone vegetated w/ Dodonaea, Sophora, Myoporum, Ohia; adj. older cone older & w/koa. Preserved_Specimen Stemmermann, R.L. 664 Hawaii BISH 1974-11-01
102 Santalum paniculatum Humuula, SE slopes of Mauna Kea; 0.5-7 km SE of Puu Kanakaleonui at treeline dry mamane woodland with mixed grass, native shrub understory Preserved_Specimen Warshauer, F.R. 3121 Hawaii BISH 1981-01-20
103 Santalum paniculatum Ocean View Estates on aa; with Metrosideros, Canthium, mamaki Preserved_Specimen Stemmermann, R.L. 697 Hawaii BISH 1975-01-10
104 Santalum paniculatum SW flank of Mauna Loa, Manuka Metrosideros Kipuka, with Coprosma, Dodonaea, Pelea, Pittosporum, Santalum, Styphelia, Vaccinium, on cinder Preserved_Specimen Higashino, P.K. 5814 Hawaii BISH 1977-06-13
105 Santalum paniculatum Kapua rough lava. Preserved_Specimen MacDaniels, L.H. 271 Hawaii BISH 1926-11-16
106 Santalum paniculatum Between Na Puukulua and 1859 lava flow dry plain Preserved_Specimen Greenwell, A.B.H. 19973 Hawaii BISH 1949-08-21
107 Santalum paniculatum National Park, Kilauea, Bird Park open woods Preserved_Specimen St.John, H. 11250 Hawaii BISH 1931-12-22
108 Santalum paniculatum Saddle Rd., small side road leading mauka from pole 116 on alluvum Preserved_Specimen Stemmermann, R.L. 676 Hawaii BISH 1975-01-08
109 Santalum paniculatum Keonepoko Iki, halfway along Kikou St. (from Kehakai Blvd.) Uluhe-covered margin of agricultural lot. Preserved_Specimen Takeuchi, W.N. 3450 Hawaii BISH 1987-02-09
110 Santalum paniculatum Pohakuloa Training Area Area 2X3 ft. Growing in pahoehoe with Pennisetum setaceum, Cocculus. Preserved_Specimen Davis, J. 296 Hawaii BISH 1980-06-07
111 Santalum paniculatum Ocean View Estates, Coral Parkway just off Tike Lane Metrosideros, Sophora, Osteomeles Preserved_Specimen Stemmermann, R.L. 699 Hawaii BISH 1975-01-10
112 Santalum paniculatum 1 mi SW of Puuanahulu, North Kona, below road sparse forest on dry aa lava Preserved_Specimen St.John, H. 22553 Hawaii BISH 1947-01-04
113 Santalum paniculatum Koaia Tree Santuary, 5 mi nw of Waimea Dry forest. Preserved_Specimen Little Jr., E.L. 31086 Hawaii BISH 1976-08-18
114 Santalum paniculatum Kau desert Preserved_Specimen Russ, G.W. Hawaii BISH 1927-04-01
115 Santalum paniculatum Kapua, makai of road, S. Kona Diospyros forest Preserved_Specimen Stemmermann, R.L. 695 Hawaii BISH 1975-11-10
116 Santalum paniculatum 20 mi from Kona aa desert Preserved_Specimen Hawaii BISH 1926-08-20
117 Santalum paniculatum SW flank of Mauna Loa, Puu Ohohia semi closed Metrosideros, Acacia forest; with Dodonaea, Styphelia, Vaccinium, on cinder Preserved_Specimen Higashino, P.K. 6331 Hawaii BISH 1977-08-05
118 Santalum paniculatum South Kohala Distr., near Puu Heewai, ca. 2 mi southeast of Keamuku Growing on edge of small gully near Pu`u He`ewai. Preserved_Specimen Davis, J. 655 Hawaii BISH 1981-12-03
119 Santalum paniculatum Puna, upper Kealakomo steep slopes of pali, lehua forest Preserved_Specimen Stone, B.C. 3018 Hawaii BISH 1959-07-09
120 Santalum paniculatum Kau desert Preserved_Specimen Forbes, C.N. 389 Hawaii BISH 1911-08-02
121 Santalum paniculatum National Park, Kipuka Puaulu between fenced area and picnic area closed canopy ohia-koa forest and exotic grass understory Preserved_Specimen Stemmermann, R.L. 666 Hawaii BISH 1974-11-01
122 Santalum paniculatum Honomalino old lava, with Lantana and Cassia glauca Preserved_Specimen Bryan Jr., E.H. 759 Hawaii BISH 1932-11-26
123 Santalum paniculatum Northeast of Pakoa Waterhole, alongside a jeep trail, Kapapala Forest Reserve, Kau District On pahoehoe substrate with ash pockets, growing in an open scattered Metrosideros scrub with Dodonaea, Styphelia, Andropogon, and Machaerina Preserved_Specimen Clarke, G. 617 Hawaii BISH 1981-07-02
124 Santalum paniculatum Kau, SE of Pahale desert Preserved_Specimen Skottsberg, C.J.F. 606 Hawaii BISH 1922-09-20
125 Santalum paniculatum S slope of Mauna Kea between Puu Kole, Puu Loaloa and Keanakolu Rd full sun; dry mixed grass, native shrub with mamane trees Preserved_Specimen Jacobi, J.D. 2005 Hawaii BISH 1981-01-20
126 Santalum paniculatum down 200 m down Hilina Pali Rd. from intersection with Chain of Craters Rd.; 20 m W of road; plot 1 Metrosideros, native shrub, Andropogon Preserved_Specimen Newell, C.L. 402 Hawaii BISH 1966-08-15
127 Santalum paniculatum Between Waha Pele and Puu Ikaaka, Hualalai, N. Kona tall shrub Preserved_Specimen Fosberg, F.R. 41649 Hawaii BISH 1961-03-22
128 Santalum paniculatum HVNP on unweathered pahoehoe scattered individuals, most in flower but no fruits Preserved_Specimen Takeuchi, W.N. 749 Hawaii BISH 1984-05-12
129 Santalum paniculatum Kaupulehu, on transect T12 Open Diospyros forest. Preserved_Specimen Takeuchi, W.N. 5810 Hawaii BISH 1989-05-20
130 Santalum paniculatum S of Papaloa, Capt. Cook, Kona open forest Preserved_Specimen Degener, O. 20342 Hawaii BISH 1949-08-13
131 Santalum paniculatum Puu Nene, near Humuula Sheep station base of small cinder cone Preserved_Specimen Herbst, D.R. 916 Hawaii BISH 1968-02-13
132 Santalum paniculatum Keauhou Ranch, Kau District Growing in disturbed open Metrosideros forest with few Acacia koa, shrubs Dodonaea, Styphelia, and pasture of Eragrostis brownei, Setaria geniculata and Sacciolepis indica. Preserved_Specimen Davis, J. 402 Hawaii BISH 1980-10-16
133 Santalum paniculatum S. Kona, N of Kahaluu, plains of Umi with Exocarpos, mamane, naio Preserved_Specimen Rock, J.F.C. Hawaii BISH 1958-04-01
134 Santalum paniculatum Above Kipuka Ahiu forest to L of 1880-1 flow Preserved_Specimen Forbes, C.N. 907 Hawaii BISH 1915-06-23
135 Santalum paniculatum N. Kona, Puuwaawaa, N slope of Hualalai, 1 km E of Kileo and Potato Hill full sun; dry koa, ohia; native shrub, exotic grass Preserved_Specimen Jacobi, J.D. 1163 Hawaii BISH 1978-08-09
136 Santalum paniculatum Kaupulehu Forest Reserve, Section B. Open bog, dry a'a', native forest at 1950 ft. Preserved_Specimen Herbst, D.R. 3004 Hawaii BISH 1973-05-03
137 Santalum paniculatum National Park, Kilauea Military Camp open Metrosideros, grassland, on unweathered pahoehoe Preserved_Specimen Takeuchi, W.N. 10 Hawaii BISH 1984-05-12
138 Santalum paniculatum N. Kona, Puuwaawaa, between Poohohoo and water tank half a mile + east of it Preserved_Specimen Kondo, Y. 77 Hawaii BISH 1946-07-20
139 Santalum paniculatum Puu Huluhulu-saddle road Preserved_Specimen Stemmermann, R.L. 665 Hawaii BISH 1974-11-01
140 Santalum paniculatum Puna; Kalapana trail Preserved_Specimen Stone, B.C. 2937 Hawaii BISH 1959-06-24
141 Santalum paniculatum N. Kona, near Huehue, main road Preserved_Specimen Cranwell, L.M. 3218 Hawaii BISH 1938-09-11
142 Santalum paniculatum Puuwaawaa Preserved_Specimen Rock, J.F.C. 2700 Hawaii BISH 1953-02-01
143 Santalum paniculatum Puu Nene, Humuula Sheep Station Preserved_Specimen Herbst, D.R. 916 Hawaii BISH 1968-02-13
144 Santalum paniculatum Puuwaawaa Preserved_Specimen Forbes, C.N. 15 Hawaii BISH 1911-06-08
145 Santalum paniculatum Kapua, S. Kona Preserved_Specimen Rock, J.F.C. 12515 Hawaii BISH 1912-06-01
146 Santalum paniculatum along Kau-Kona road Preserved_Specimen Skottsberg, C.J.F. 607 Hawaii BISH 1922-09-21
147 Santalum paniculatum 9 mi W of Naalehu Post Office on Hwy. 11 Preserved_Specimen Wagner, W.L. 5233 Hawaii BISH 1984-01-16
148 Santalum paniculatum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 17149 Hawaii BISH 1918-07-01
149 Santalum paniculatum off the North Kohala Rd. at Acacia koaia Sanctuary, Dept. of Forestry Preserved_Specimen Wagner, W.L. 5285 Hawaii BISH 1984-02-28
150 Santalum paniculatum Puuwaawaa, N. Kona, 30.8 mi N of intersection of Kealakikua Rd. and Belt Rd Preserved_Specimen Ozaki, E.T. 1337 Hawaii BISH 1954-12-28
151 Santalum paniculatum Kapua, S. Kona Preserved_Specimen Rock, J.F.C. 12515 Hawaii BISH 1912-01-01
152 Santalum paniculatum Kilauea Preserved_Specimen Bryan, W.A. 634 Hawaii BISH 1902-10-01
153 Santalum paniculatum S slope Mauna Kea Preserved_Specimen Skottsberg, C.J.F. 5 Hawaii BISH 1935-08-02
154 Santalum paniculatum Kilauea Preserved_Specimen Rock, J.F.C. 8770 Hawaii BISH 1911-08-01
155 Santalum paniculatum N. Kona, Hawaii Belt Rd., near mile 28, Kaupulehu Preserve Preserved_Specimen Flynn, T.W. 1302 Hawaii BISH 1985-10-16
156 Santalum paniculatum Volcano of Kilauea Preserved_Specimen Rock, J.F.C. 8770 Hawaii BISH 1911-08-01
157 Santalum paniculatum Kilauea Volcano Preserved_Specimen Giffard, W.M. 8000 Hawaii BISH 1915-10-01
158 Santalum paniculatum National Park, Chain of Craters Rd. ~3 mi from jct. with Crater Rim Rd Preserved_Specimen Sohmer, S.H. 6336 Hawaii BISH 1970-01-08
159 Santalum paniculatum National Park, Makaopuhi-Kalapana trail Preserved_Specimen Stone, B.C. 2937 Hawaii BISH 1959-06-24
160 Santalum paniculatum Ocean View Estates, 0.1 mi N. of Ocean View General Store, along Hwy. 11 Preserved_Specimen Kimura, B. 5 Hawaii BISH 1980-06-29
161 Santalum paniculatum Kilauea, Bird Park; Hawaii Natl Park open woods Preserved_Specimen St.John, H. 11265 Hawaii BISH 1931-12-22
162 Santalum paniculatum Pohakuloa Training Area. Growing on small gulch mauka of telephone pole #116 Preserved_Specimen Stemmermann, R.L. 678 Hawaii BISH 1975-01-08
163 Santalum paniculatum Between Huehue and Puuwaawaa Preserved_Specimen Rock, J.F.C. 3613 Hawaii BISH 1909-06-01
164 Santalum paniculatum Kaalapunuwale, Kona Preserved_Specimen Forbes, C.N. 228 Hawaii BISH 1911-06-21
165 Santalum paniculatum Slopes of Mauna Kea, Waiki Preserved_Specimen Forbes, C.N. 460 Hawaii BISH 1911-08-01
166 Santalum paniculatum S. Kona, Puu Leihua Preserved_Specimen Rock, J.F.C. 25728 Hawaii BISH 1957-03-16
167 Santalum paniculatum In gulch with Lantana-Ipomoea congesta association. Land of Kapua, S. Kona District Preserved_Specimen Nagata, K.M. 2609 Hawaii BISH 1983-02-13
168 Santalum paniculatum open woods, Aleale Crater, chain of craters, Napau Trail Kilauea Preserved_Specimen St.John, H. 11222 Hawaii BISH 1931-12-20
169 Santalum paniculatum Makaopuhi-Kalapana trail Preserved_Specimen Stone, B.C. 2940 Hawaii BISH 1959-06-24
170 Santalum paniculatum Puu Nene, near Humuula Sheep Station Preserved_Specimen Herbst, D.R. 916 Hawaii BISH 1968-02-13
171 Santalum paniculatum Kilauea Volcano Preserved_Specimen Giffard, W.M. 8000 Hawaii BISH 1915-10-01
172 Santalum paniculatum Puuwaawaa Preserved_Specimen Ozaki, E.T. 1375 Hawaii BISH 1954-12-31
173 Santalum paniculatum N.Kona, Puuwaawaa; along Huehue Puuwaawaa pipe line, between water tanks 1-3 Preserved_Specimen Kondo, Y. 111 Hawaii BISH 1946-07-24
174 Santalum paniculatum National Park, Makaopuhi-Kalapana trail Preserved_Specimen Stone, B.C. 2940 Hawaii BISH 1959-06-24
175 Santalum paniculatum n. Kona district. Kaupulehu Preserve, along Mamahaloa Highway (190) at 12 miles NE of Kailua Kona (by 28 mile marker); Preserved_Specimen Lorence, D.H. 5484 Hawaii BISH 1987-08-06
176 Santalum paniculatum Puuwaawaa, N. Kona, 30.8 mi N of intersection of Kealakikua Rd. and Belt Rd Preserved_Specimen Ozaki, E.T. 1338 Hawaii BISH 1954-12-28
177 Santalum paniculatum Puuwaawaa Preserved_Specimen Skottsberg, C.J.F. 686 Hawaii BISH 1922-09-26
178 Santalum paniculatum Kilauea Volcano Preserved_Specimen Giffard, W.M. 12517 Hawaii BISH 1915-10-30
179 Santalum paniculatum Kanehaha, Kona Preserved_Specimen Forbes, C.N. 236 Hawaii BISH 1911-06-23
180 Santalum paniculatum Manuka Mauka, Kau Preserved_Specimen St.John, H. 11340 Hawaii BISH 1931-12-26
181 Santalum paniculatum Puhulahula, hill in Humuula Saddle 20 mi w. of Hilo. Acacia koa forest. Mauna Loa Forest and Game reserve Preserved_Specimen Little Jr., E.L. 31264 Hawaii BISH 1976-09-23
182 Santalum paniculatum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 13047 Hawaii BISH 1917-08-01
183 Santalum paniculatum Pohakuloa military reservation. Open metrosideros, myrsine and Dodonea on substation Preserved_Specimen Higashino, P.K. 5233 Hawaii BISH 1977-01-13
184 Santalum paniculatum Puuwaawaa Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-28
185 Santalum paniculatum North Kona, Puuwaawaa Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-28
186 Santalum paniculatum Kilauea Crater Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
187 Santalum paniculatum tree in brown Ranch pasture Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-21
188 Santalum paniculatum Mauna Kea Preserved_Specimen Bryan, L.W. Hawaii BISH 1951-01-19
189 Santalum paniculatum Old stream crack, Kilauea Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-21
190 Santalum paniculatum Hawaii Preserved_Specimen Adee, K.T. Hawaii BISH
191 Santalum paniculatum near Kilauea Preserved_Specimen Rock, J.F.C. Hawaii BISH 1958-10-06
192 Santalum paniculatum N. Kona, lava fields, Kaupulehu Preserved_Specimen Bryan, L.W. Hawaii BISH 1959-09-09
193 Santalum paniculatum Kilauea, trail to crater from volcano house Preserved_Specimen Neal, M.C. Hawaii BISH 1927-08-03
194 Santalum paniculatum Hawaii Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-23
195 Santalum paniculatum Kilauea, in old stream creek Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-06-21
196 Santalum paniculatum N.Kona, Puuwaawaa, 4/5 of way east of tank behind Poohohoo, near and in sight of Halfway House Preserved_Specimen Kondo, Y. Hawaii BISH 1946-07-18
197 Santalum paniculatum Kilauea Volcano Preserved_Specimen Rock, J.F.C. Hawaii BISH 1911-07-01
198 Santalum paniculatum Hawaii National Park, Kilauea, Kau Desert in the vicinity of great crack Preserved_Specimen Judd Jr., A.F. Hawaii BISH 1939-03-11
199 Santalum paniculatum Kilauea Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-21
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